Module 8

Global Comparison

A quantitative synthesis across all biomes — net primary productivity, area, carbon storage, biodiversity, and climate-change exposure — in a single reference view. Values are representative; see per-biome modules for full sourcing.

1. NPP, Area & Carbon Stock

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2. Comprehensive Summary Table

BiomeArea (M km2)MAT (°C)MAP (mm)NPPSoilVegetation
Tropical rainforest~1725–282000–40002200Oxisol / UltisolBroadleaf evergreen
Temperate forest~115–15750–1500600–800Alfisol / MollisolDeciduous hardwoods
Grassland / savanna~5210–28300–1500400–900Mollisol / VertisolGrasses, forbs
Desert~5520–35<25045AridisolSucculents, annuals
Boreal forest~17−10–5300–900360Spodosol / HistosolConifer trees
Tundra~8−25–550–250140GelisolMosses, sedges, shrubs
Mediterranean~2.510–20300–700500–800Alfisol / EntisolSclerophyllous shrubs
Freshwater~2VariableVariable250AquicAquatic macrophytes
Coral reef~0.623–29 (water)2500Calcium carbonateScleractinian corals
Open ocean~332−2–30 (water)125Phytoplankton

NPP in g C m-2 yr-1.

3. Climate-Change Projections (RCP8.5 by 2100)

  • Tropical forest: increased drought frequency; Amazon dieback risk in southeast arc (Nobre 2016).
  • Temperate forest: poleward range shifts up to ~500 km; increased fire risk; species-range mismatches with interacting pollinators.
  • Grassland / savanna: bush encroachment in many regions; altered fire regimes; C4-to-C3 shifts under elevated CO2.
  • Desert: expansion 4–11%; increased dust storms; Sahelian greening (unusually) with higher CO2.
  • Boreal forest: northward expansion (~5–8 km decade-1); insect outbreaks; severe wildfire seasons.
  • Tundra: shrubification; 30–60% area reduction by 2100; permafrost-C release.
  • Aquatic: coral bleaching every 2–3 yr globally; ocean acidification; deoxygenation; polar ocean warming 3°C+.

4. Conservation Status (WWF Global 200)

The WWF’s Global 200 analysis (Olson 2001) identified 233 ecoregions of exceptional biodiversity value across the world’s biomes. Of these, ~73% are either critically endangered, endangered, or vulnerable. Highest-threat biomes: Mediterranean forests (>90% converted), temperate broadleaf forests, and tropical dry forests. Lowest-threat biomes by conversion: tundra, boreal forest (but rapidly warming), deep pelagic.

5. Course Synthesis

Eight modules traced Earth’s major biomes from tropical rainforest through temperate forest, grassland, desert, boreal forest, tundra, and aquatic systems back to a comparative synthesis. The overarching pattern is that two climate variables — temperature and precipitation — predict biome identity worldwide, while secondary drivers (fire, soil, biogeographic history) refine the picture. Every biome is now being reshaped by anthropogenic pressure, most acutely climate change and land-use. Understanding biomes is the prerequisite for understanding what a future Earth’s biosphere will look like.

Further Reading

• Whittaker, R. H. (1975). Communities and Ecosystems. Macmillan.

• Chapin, F. S., Matson, P. A. & Vitousek, P. M. (2011). Principles of Terrestrial Ecosystem Ecology, 2nd ed. Springer.

• Ricklefs, R. E. & Relyea, R. (2019). The Economy of Nature, 8th ed. Freeman.

• IPCC AR6 Working Group II (2022). “Terrestrial and freshwater ecosystems and their services.” Chapter 2.

• Olson, D. M. et al. (2001). “Terrestrial ecoregions of the world.” BioScience, 51, 933–938.

• Schimel, D. S. (1995). “Terrestrial biogeochemical cycles: global estimates with remote sensing.” Remote Sens. Environ., 51, 49–56.

• Bond, W. J. (2019). Open Ecosystems: Ecology and Evolution Beyond the Forest Edge. Oxford UP.

• Field, C. B. et al. (1998). “Primary production of the biosphere: integrating terrestrial and oceanic components.” Science, 281, 237–240.